The importance of animal pollinators to agricultural productivity and as a component of biodiversity is well-understood (Klein et al., 2007; Ollerton et al., 2011). However, declines in the abundance and distribution of many pollinator species have prompted concerns for the maintenance of their pollination services (e.g. Goulson et al., 2015; Bartomeus et al., 2019). While several causes have been identified, there is a general consensus that agricultural intensification is a key factor (Ollerton et al., 2014; Goulson et al., 2015; Balfour et al., 2018).